B.S., Biology, University of Puerto Rico, 1976.
M.S., Zoology, University of Idaho, 1979.
Ph.D., Animal Ecology, Iowa State University, 1985.
Status and population ecology of the brown pelican, Pelecanus occidentalis, in the Greater Puerto Rican Bank. Ph.D. Dissertation, Department of Animal Ecology, Iowa State University, 1985.
Breeding biology and food habits of the great blue heron at Heyburn State Park, Benewah County, Idaho. M.S. Thesis