Godwin, J.R. and D.F. Fautin. 1992. Defense of host actinians by anemonefishes.
Copeia 1992(3): 902-908.

Entacmaea quadricolor is the most common and geographically widespread host anemone. In this paper, we present strong support for the hypothesis that the anemonefish symbionts of E. quadricolor provide aggressive protection against anemone predators and assert that this symbiosis is therefore mutualistic. Observations suggesting that certain of the symbioses between other anemonefishes and actinians are also mutualistic are presented.